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The physiology of the plant-mycorrhizal.fungus interface: preliminary observations

poster
AYLING. S.M., S.E.SMITH & F.A.SMITH
Department of Soil Science, Waite Research Institute, University of Adelaide, Adelaide, S.A.5064, Australia

ICOM1 Abstract
The symbiosis between a host plant and mycorrhizal fungus involves bidirectional transfer of nutrients. Numerous workers have demonstrated enhanced uptake of phosphorous from the soil to the plant in the presence of mycorrhizal fungi; and export of carbon metabolites from the plant to the fungal hyphae. The mechanisms controlling these transfers must involve some modification of both plant and fungal membranes. There is no information about the physiological conditions in the region of contact between the two organisms. Histochemical techniques have demonstrated high ATPase activity in the periarbuscular membrane, while molecular biological methods have identified a phosphate transporter in a VAM fungus. In the absence of detailed knowledge of the membrane potentials and ionic gradients in this zone the functional behaviour of these proteins can only be assumed. In this study we are combining electrophysiological techniques and confocal laser.scanning microscopy to characterise the host-fungus interface.