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The use of phytate as a phosphorus source by mycorrhizal and non-mycorrhizal Scots pine (Pinus sylvestris)

talk
session 1: physiology
COLPAERT JAN V.1, ANDRÈ VAN LAERE1 & KATIA K. VAN TICHELEN2
1Laboratory of Developmental Biology,
2Laboratory of Plant Ecology, Institute of Botany, Katholieke Universiteit Leuven, Kardinaal Mercierlaan 92, B-3001 Leuven, Belgium

ICOM1 Abstract
The use of phytate-P by mycorrhizal and non-mycorrhizal pine seedlings was investigated. Mycorrhizal plants were either inoculated with Thelephora terrestris or Suillus luteus. Plants were grown for 9 weeks in a semi-hydroponic system with P as the growth-limiting factor. Addition of a relatively high concentration of soluble Ca-phytate resulted in an increased relative growth rate (RGR) in both mycorrhizal and non-mycorrhizal plants. The effect of the mycorrhizal fungi on the use of the phytate-P was small. The addition of an insoluble phytate to the growth substratum did not result in an increase of the RGR and P uptake neither in the non-mycorrhizal, nor in the mycorrhizal pines. Soil phytate, which has a low solubility, is probably not a very useful P source for Thelephora terrestris and Suillus luteus. Nevertheless the external mycelia of these fungi exhibited a substantial extracellular acid phosphatase activity. The activity was correlated with the fungal biomass in the growth substratum and decreased with an increasing P nutrition. Phytase activity of the external mycelia could not be detected although a low phytase activity could be observed at the root surface. Mycorrhizas showed significantly higher activities than uninfected roots. The orthophosphate, labile and stable P fractions in the substratum were analysed at harvest. In the mycorrhizal treatments the Pi pool decreased significantly whereas the more stable P pool tended to increase.