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Ca:Al ratio effects on competition between arbuscular mycorrhizal and ectomycorrhizal forest trees

POSTER OR TALK: Biodiversity and Ecology or Mycorrhizae in Stressed Ecosystems
DECKER, KELLY L.M, SHERRI J. MORRIS, and RALPH E.J. BOERNER
Department of Plant Biology, Ohio State University, Columbus, OH 43210 USA

ICOM1 Abstract
In the deciduous forest of eastern North America, the abundance of species of trees dependent on ectomycorrhizae, such as oaks (Quercus spp.) and hickories (Carya spp.) has declined over the last two decades while the abundance of species dependent on arbuscular mycorrhizae, such as maples (Acer spp.) and yellow-poplar (Liriodendron tulipifera) has increased. We hypothesize that this shift in dominance is the consequence of the effects of heavy N deposition and decreasing soil Ca:Al ratio on both plants and fungi. In this study, we examine the effect of lowering the Ca:Al ratio from 100 to 4 under high N conditions on growth and competitive interactions between second year seedlings of ectomycorrhizal red oak (Quercus rubra) and arbuscular mycorrhizal yellow-poplar (Liriodendron tulipifera). Overall, the relative growth rate of yellow-poplar seedlings exceeded that of red oak seedlings, although the difference was greater at the higher Ca:Al ratio. When in interspecific competition, yellow-poplars outgrew red oaks at both Ca:Al ratios. The relative growth rate of red oak seedlings was relatively insensitive to competition, whereas yellow-poplar plants grew faster when in inter- than intraspecific competition. Under these high N conditions, yellow-poplars were competitively superior to red oaks regardless of Ca:Al ratio. We will also discuss the impact of Ca:Al ratio on N uptake, C and N allocation to leaves, stems, and roots and the effects of Al on the fine structure of ectomycorrhizal roots.